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Gravitas and the members of the Chamber of the Ancients have spoken. It has been determined that the fate of the Autobots will rest upon the shoulders of an archivist named Optronix. Although Optronix is initially reluctant to accept this great responsibility, he begins to understand the devastating risks at stake for the entire Transformers race.

Optronix is given the leadership name of Optimus Prime once the power of The Matrix is bestowed onto him. It is a sign that he must lead the the Autobots away from Cybertron, not only to escape the war with the Decepticons, but also to escape the self-destructive war that exists within them. After a colossal battle with Megatron that spans the boundaries of both time and space, Optimus discovers the power within himself to step up and provide the leadership is race needs in their most desparate hour.